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The C.A.S.H. Courier
ARTICLE from the Winter 2009 Issue
The Ineffectiveness of Coyote Hunting
Dr. Robert Crabtree, President and Founder of YELLOWSTONE ECOSYSTEM
STUDIES
Biology Department, Montana State University, answers the question “What
effect does reduction of coyotes (older than 6 months) have on the remaining
population?” He had heard claims that reduction of adult coyotes would
lessen predation on domestic sheep or game animals such as mule deer or
antelope. He refutes those claims with the following points:
Most reduction programs are indiscriminate in nature, meaning the
individuals removed (killed) are probably not the offending individuals.
Even if some offending individuals are removed, there is great likelihood
that the responses described below will take place anyway. Although removal
of offending individuals can temporarily alleviate predation rates, the
alleviation is usually short-term and likely has long-term side-effects that
make control activities ineffective. It can not be over-emphasized how
powerfully coyote populations compensate for population reductions. Both
evolutionary biology and the results of my research the last three years
indicate that the basis of this resiliency is embedded in the evolutionary
past of the coyote.
(1) Actual reduction in the density and number of coyotes does occur but
is compensated by immediate immigration into the reduction area by lone
animals or shifts in surrounding social groups.
(2) Reduction results in a smaller social group size which increases the
food per coyote ratio. This food surplus is biologically transformed into
higher litter sizes and higher litter survival rates. The increase in food
availability improves the nutritional condition of breeding females which
translates in higher pup birth weights and higher pup survival.
(3) Reduction causing higher pup survival is fundamentally a function of
the general mammalian reproductive strategy that delays the majority of
reproductive energetic investment beyond the gestation period, e.g., young
pups. Thus, the normal litter of six pups has a good chance of (a.)
surviving the typically high summer mortality period and, (b.) being
recruited into the pack the following winter as adults.
(4) Reductions of adult-sized coyotes 6 months or older results in
smaller pack size, which leaves fewer adults to feed pups. This may further
add incentive for the remaining adults to kill larger prey as well as
putting pressure on the adults to select for the most vulnerable prey close
to areas of human activity. Because predators like coyotes also learn what
is appropriate food when they are pups, and are reluctant to try ‘new’ food
sources unless under great stress (such as having to feed a large litter of
pups) reduction programs, in effect, may be seen as forcing coyotes to try
new behaviors (eating domestic livestock) which they would otherwise avoid.
(5) Reductions cause an increase in the percentage of females breeding.
(6) Reduction causes the coyote population structure to be maintained in
a colonizing state. For example, the average age of a breeding adult in an
unexploited population is 4 years old. By age 6 reproduction declines,
whereby older, alpha pairs maintain territories but fail to reproduce. This
may eliminate the need to kill sheep or fawns in the early summer in order
to feed pups. Exploiting or consistently reducing coyote populations keeps
the age structure skewed to the young and in a state of constant social and
spatial flux. Therefore, the natural limitations seen in older populations
are absent and the territorial, younger populations are much more
productive.
(7) Reductions cause young adults (otherwise prone to dispersing) to stay
and secure breeding positions in the exploited area. This phenomenon is
well-documented. There are other demographic responses that negate the
effectiveness of control practices but the aforementioned covers the most
important.
Alternate prey The preferred, natural prey of coyotes is rodents, such as
voles and jackrabbits, animals that require ample cover and supplies of
forage. Because domestic livestock grazing has been increased on many
western range lands, grasses and protective cover have been greatly reduced,
leaving coyotes with fewer preferred prey. Coyotes only switch to
alternative prey such as livestock when a preferred prey item is absent or
in low numbers. Current grazing practices may therefore be seen as a cause
of predation on other species like domestic sheep
We do not yet know how coyote populations will respond once predator
reduction programs are terminated. We should ask, “Are there other
non-lethal management alternatives that may be effective in reducing
predation on domestic livestock?” In conclusion, the common practice of
reducing adult coyote populations on western rangelands is most likely
ineffective and may even increase the number of lambs, fawns, and calves
killed by coyotes.
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